IN ORDER OF IMPORTANCE:
Alderfox and Hazelthorn, equally important so they go together! Their Beetle's brothers, with Alder being older and Hazel being younger. They have similar personalities, so similar that they seem to always be at odds! Both of them are competitive and ambitious, always looking to outdo the other. The only thing they've ever been united in is protecting Beetleshine's precious light.
Willowbloom, the mother! Willowbloom had been seduced by the loner Quintley, who disappeared when she told him she was pregnant. She begged her best friend, Foxtail, to claim the kits as his own. She'd kept the secret ever since. Whether she genuinely fell in love with Foxtail is up to you!
Foxtail, the adoptive father! The only ones who know he isn't their birth father is him and Willowbloom. He'd been in love with his best friend for moons, and gladly took the role as father of her kits when the time came. Whether he's still in love with her, fell out of love, or even told her at all, is up to you!